Reaction-diffusion equations are a trusted modeling framework for the dynamics of biological populations in space and time, and their traveling wave solutions are interpreted as the density of an invasive species that spreads at constant speed. Even though certain species can significantly alter their abiotic environment for their benefit, and even though some of these so-called "ecosystem engineers" are among the most destructive invasive species, most models neglect this feedback. Here, we extended earlier work that studied traveling waves of ecosystem engineers with a logistic growth function to study the existence of traveling waves in the presence of a strong Allee effect. Our model consisted of suitable and unsuitable habitat, each a semi-infinite interval, separated by a moving interface. The speed of this boundary depended on the engineering activity of the species. On each of the intervals, we had a reaction–diffusion equation for the population density, and at the interface, we had matching conditions for density and flux. We used phase-plane analysis to detect and classify several qualitatively different types of traveling waves, most of which have previously not been described. We gave conditions for their existence for different biological scenarios of how individuals alter their abiotic environment. As an intermediate step, we studied the existence of traveling waves in a so-called "moving habitat model", which can be interpreted as a model for the effects of climate change on the spatial dynamics of populations.
Citation: Maryam Basiri, Frithjof Lutscher, Abbas Moameni. Traveling waves in a free boundary problem for the spread of ecosystem engineers[J]. Mathematical Biosciences and Engineering, 2025, 22(1): 152-184. doi: 10.3934/mbe.2025008
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Reaction-diffusion equations are a trusted modeling framework for the dynamics of biological populations in space and time, and their traveling wave solutions are interpreted as the density of an invasive species that spreads at constant speed. Even though certain species can significantly alter their abiotic environment for their benefit, and even though some of these so-called "ecosystem engineers" are among the most destructive invasive species, most models neglect this feedback. Here, we extended earlier work that studied traveling waves of ecosystem engineers with a logistic growth function to study the existence of traveling waves in the presence of a strong Allee effect. Our model consisted of suitable and unsuitable habitat, each a semi-infinite interval, separated by a moving interface. The speed of this boundary depended on the engineering activity of the species. On each of the intervals, we had a reaction–diffusion equation for the population density, and at the interface, we had matching conditions for density and flux. We used phase-plane analysis to detect and classify several qualitatively different types of traveling waves, most of which have previously not been described. We gave conditions for their existence for different biological scenarios of how individuals alter their abiotic environment. As an intermediate step, we studied the existence of traveling waves in a so-called "moving habitat model", which can be interpreted as a model for the effects of climate change on the spatial dynamics of populations.
The Business Process Model and Notation (BPMN) [15] has been widely adopted in recent years as one of the standard languages for visual description of business processes. In particular BPMN has been extensively used in facilitating effective communication among technical experts, business users and other stakeholders within a work group, for business process design and implementation, or problem solving in business domains.
BPMN however does not contain a formal semantics. Consequently, rigorously specifying dynamical behaviors of objects in a BPMN specified business process model can be challenging and error-prone. For critical systems (e.g., a medical process specified in BPMN), the existence of semantic ambiguities in these systems is costly from every perspective and thus should be eliminated as much as possible. For these reasons, in recent years, several efforts have been made to assign BPMN with formal semantics. Among them, [3] and [4] in particular proposed to map a BPMN model into a Petri net, which is a mathematical modeling language (although a Petri net can be graphical too) that has precise execution semantics. In addition, decades of research in Petri nets offers well-established theories to support analysis of dynamic properties in Petri-net specified systems. In [3] and [4], a given BPMN model is first dissected into several BPMN elements. For each type of the element, a mapping from the element to a Petri net module is proposed. After the dissection step and mapping step, these Petri nets modules are eventually integrated together to construct a single Petri net.
In our previous work ([17], [18], [19]) meanwhile we presented a straightforward first-order-logic (FOL) based framework for Petri nets. More specifically, we have shown that: a) we can represent Petri nets and their variants as Situation-Calculus1 based action theories, and the theories for basic Petri nets are called in short as SCOPE, standing for Situation Calculus Ontology for PEtri nets ([17], [18]); b) given an instance of SCOPE theory, we could further build up Golog procedures2, where sequential, iterative, or nondeterministic composite sequences of transition firings in Petri nets and their variants can be further axiomatized through defining macros; these macros are built on top of more primitive procedures and the only basic action in the domain transition firing [19]; c) executability testing of Golog procedures via theorem-proving can be implemented through efficient logic programming in Prolog ([18], [19]).
1The Situation Calculus ([10,16]) is a particular formal language for describing changes upon actions in dynamical domains.
2In the Situation Calculus, a Golog procedure defines a complex action that is composed of basic actions.
In this paper, we show that the mapping method as proposed in ([3] and [4]) can be naturally extended into SCOPE context. That is, using SCOPE, we can easily define a Petri net module, resulted from performing mapping on a BPMN object, as a Golog complex procedure. These Golog procedures act as interfaces encapsulating Petri net from being directly accessed by end users. Consequently, stakeholders can still work at BPMN level, while the model is enriched with a Petri-net specified formal semantics. Additionally, with these procedures, users can write more complicated user-defined BPMN procedures in a hierarchical way. Since a BPMN procedure can be further translated into Prolog programs [19], our approach support automated verification of important system dynamic properties, such as the executability of these procedures.
The remainder of this paper is organized as follows. Section 2 provides introductory reviews to the key technologies employed in this paper. Enriched with an example, Section 3 is the core section that explains our approach. Conclusive remarks are presented in Section 4.
In this section, we give a brief introduction to BPMN and Petri Nets. The method of mapping from BPMN onto Petri Nets as stated in Section 3 of [4] is then briefly reviewed. We also introduce here Situation Calculus/Golog and SCOPE.
Business Process Model and Notation (BPMN) is a notational system for graphical specification of processes in business models. BPMN was developed and has been maintained by the Object Management Group (OMG) [15]. Control-flow related elements in BPMN are the focus of this paper and they include objects of events, tasks and gateways.
An event denotes happening of something in a model and is represented as a circle. In particular, a start/end event signals the start/finish of a process. An activity denotes work to be done in a model and is represented as a rounded-corner rectangle. A task is an atomic activity. Gateways are diamond-shaped quadrilaterals and they denote different routing constructs. Among them, a fork gateway (AND-split) creates concurrent flows and a join gateway synchronises these flows. A decision gateway (XOR-split) selects exactly one flow from a set of them based on certain criteria (data or event) and a merge gateway (XOR-join) merges the alternative flows in the set into one flow. When the choice is non-unique, the pair of gateways OR-split and OR-join is applied instead. Graphically these BPMN objects are connected to each other by arrows. Figure 2 is an example BPMN-denoted business process (adapted from Fig. 13 (a) in [4]).
A Petri net (PN) [14] is a pair
The method of mapping a BPMN process into a Petri net introduced in [4] assumes that the BPMN process is in the first place "well-formed". A well-formed BPMN should satisfy several conditions (e.g., a start event has one outgoing flow and no incoming flow.). It is shown that a given BPMN process can always be transformed into a well-formed one ([1,9,25,26]). The mapping process by itself is straightforward and are summarized in Figure 1 (Several issues are identified and addressed accordingly in [4]; however they are not discussed here). In the figure, note that places in the Petri net modules/components are dashed, meaning that these places would act as connectors to link multiple components into a Petri net through mapping. As an example, the Petri net in Figure 3 is resulted from performing mapping on the BPMN model in Figure 2.
The Situation Calculus is a logical language for representing actions and changes in a dynamical domain. It is first proposed by McCarthy and Hayes in 1969 [10]. The language
The theory
Primitive Action
Fluent
Situation-Independent relations
•
•
•
The Foundational Axioms
(∀s,p,t)(Poss(fire(t),s)≡(pre(p,t)⊃Tkns(p,s)≥1)) |
defines the condition for a transition node
(∀s,p,a,n)(Tkns(p,do(a,s))=n≡γf(p,n,a,s)∨ |
(Tkns(p,s)=n∧¬∃n′γf(p,n′,a,s))), |
where
•
•
The above Successor State Axiom summarizes all conditions where the number of tokens at place
As an example, we define Petri net in Figure 3 as a SCOPE theory. The axiomatic part of SCOPE, including Foundational Axioms, Precondition Axioms, and Successor State Axioms remain unchanged. We only need to specify the initial marking and the situation-independent, topological structure of the Petri-net in Figure 3. That is, the number of tokens at place
pre(P1,T1),pre(T1,P2),…,pre(T11,P12). |
This section includes two parts. We first specify formally the restrictions for well-formed BPMN processes, this would facilitate implementation of automated tools for evaluating formally whether a BPMN process is well-formed (using a FOL theorem prover for example). We then demonstrate how Petri nets modules for their corresponding BPMN objects can be formally described through carrying out further axiomatization on top of SCOPE.
From Section 3 of [4], a well-formed BPMN process has to satisfy the following five criteria:
1. a start event or an exception event has just one outgoing (sequence) flow but no incoming flow;
2. an end event has just one incoming flow but no outgoing flow;
3. activities and intermediate events have exactly one incoming flow and one outgoing flow;
4. fork or decision gateways have one incoming flow and more than one outgoing flows; and
5. join or merge gateways have one outgoing flow and more than one incoming flows.
In this section, first-order axiomatization of these criteria are presented (assuming that objects in the domain are sorted in logical sense).
•
•
•
•
•
•
• A start/exception event has one outgoing flow and does not have incoming flow
(∀x,y)(¬(pre(x,start(y))∧¬(pre(x,exception(y))), |
(∀y)((start(y)∨exception(y))⊃((∃x)pre(y,x)∧ |
¬(∃x1,x2)(pre(y,x1)∧pre(y,x2)∧x1≠x2))). |
• An end event has one incoming flow and does not have outgoing flow
(∀x,y)¬(pre(end(y),x)). |
(∀y)(end(y)⊃((∃x)pre(x,y)∧ |
¬(∃x1,x2)(pre(x1,y)∧pre(x2,y)∧x1≠x2))). |
• An activity/intermediate-event has one and only one incoming flow and outgoing flow
(∀y)((activity(y)∨interE(y))⊃ |
((∃x)pre(y,x)∧¬(∃x1,x2)(pre(y,x1)∧pre(y,x2)∧x1≠x2)) |
((∃z)pre(z,y)∧¬(∃z1,z2)(pre(z1,y)∧pre(z2,y)∧z1≠z2))). |
• An gateway fork or decision has one incoming flow and more than one outgoing flows
(∀y)((fork(y)∨decision(y))⊃((∃x)pre(x,y)∧ |
¬(∃x1,x2)(pre(x1,y)∧pre(x2,y)∧x1≠x2))∧ |
(∃x1,x2)(pre(y,x1)∧pre(y,x2)∧x1≠x2))). |
• An gateway join or merge has one outcoming flow and more than one ingoing flows
(∀y)((join(y)∨merge(y))⊃((∃x)pre(y,x)∧ |
¬(∃x1,x2)(pre(y,x1)∧pre(y,x2)∧x1≠x2))∧ |
(∃x1,x2)(pre(x1,y)∧pre(x2,y)∧x1≠x2))). |
It is known from our previous work that, given a SCOPE theory, we can write SCOPE procedures through defining macros, where these macros are built on top of more primitive, existing procedures and the only basic action in the domain -transition node fire. With respect to the Petri net in Figure 3, we can define a set of simple SCOPE procedures. For example,
proc
endProc
proc
endProc
The procedure
The procedure
With two procedures:
proc
endProc
To this end, it is quite clear how methodologically we are able to axiomatize these BPMN flow-control components: we can simply define those Petri-net modules transformed from those BPMN objects, as illustrated in Figure 1, as SCOPE procedures.
We start from axiomatizing a Petri-net module for the start object
proc
endProc
That is, as long as the topological relationship is verified,
The definition of a procedure corresponding to a fork gate in BPMN involves one place node that enters (and multiple place nodes that leave) the transition node to be fired. Suppose that
proc
endProc
The definition of a procedure corresponding to a join gate in BPMN involves one place node that leaves (and multiple place nodes that enter) the transition node to be fired. Suppose
proc
endProc
The definition of a procedure corresponding to a decision gate in BPMN involves one place node that enters at least one transition node, where each transition node in turn enters a place node. Suppose that
proc
;
endProc
The definition of a procedure corresponding to a merge gate in BPMN involves one place node that multiple transition nodes enter, where for each transition node there is a place node that enters the node. Suppose that
proc
;
endProc
This section includes an example of an order process (adapted from Fig. 13(a) in [4]). When an order request arrives from a customer, both tasks "CheckCreditCard" and "Prepare Products" are initialized. If the credit card is OK, the packed products would be shipped (the task "ShiProducts" is executed) and the process finishes. Alternatively, if the credit card fails, the process finishes directly. BPMN representation of the process is depicted in Fig. 2.
The following sentences capture the topology of the Order Process Diagram.
The nine BPMN objects in Figure 2 are defined as the following SCOPE procedures.
proc
endProc
proc
endProc
proc
endProc
proc
endProc
proc
;
endProc
proc
endProc
proc
endProc
proc
;
endProc
proc
endProc
It is handy to create further user-defined new procedures through macro-actions, i.e., through hierarchically constructing more complex procedures from existing ones. For example, the following procedure presumably specifies all possible executions of the BPMN process as specified in Figure 2.
proc
endProc
Feasibility/executability of procedure
?−Do(orderProcess,S0,S), |
any instantiation of the situation variable
This paper proposes to axiomatize the Petri nets modules, resulted from transforming BPMN objects, as SCOPE-based procedures. In fact, the impressive feature of high adaptability and extensibility is shared by all Situation-Calculus-based Basic Action Theories including SCOPE. In a similar way, we recently used SCOPE to aggregate event patterns in Supply Chain Management [21], and to mitigate adverse interaction analysis for concurrent application of clinical practise guidelines for comorbid patients [11,20]. Ongoing approaches using directly first-order logic for integrating medical treatments to manage multimorbidity include [11,27,12,13,22]. Formal process knowledge representation and reasoning might also find its applicability in combining probabilistic graphical models to build up frameworks and systems in areas for example information retrieval [23,24].
Our previous research ([18] and [19]) indicates that we can easily implement software tools using logic programming language Prolog [2]. Further, we might consider adopting tractable variant to Situation Calculus (for example, a sequence of research on the topic proposed in [6,7,5] is quite interesting and relevant) so an actual implementation became practically feasible. Given the rapid development of first-order theorem proving technologies in the recent years, theorem provers have recently been used to support reasoning in ontologies [8]. As part of our initiative, we plan to adapt theorem proving technologies for process ontologies like SCOPE theories.
Finally, we remark that our approach is able to incorporate processes specified with temporal constraints, since in our previous research reported in [19], a variant to SCOPE which axiomatize Time Petri-nets are presented.
We are thankful to the insightful reviews we have received to improve both the content and presentation of the paper. We gratefully acknowledge the support by NSERC (Natural Sciences and Engineering Research Council of Canada) CREATE (Collaborative Research and Training Experience Program) award in ADERSIM (Advanced Disaster, Emergency and Rapid-response Simulation)3, ORF-RE (Ontario Research Fund -Research Excellence)4 award in BRAIN Alliance5, Dapasoft Inc. in Toronto, ON, Canada6, and the York Research Chairs Program.
3http://www.yorku.ca/adersim/NSERC/
4https://www.ontario.ca/page/ontario-research-fund-research-excellence
[1] | A. N. Kolmogorov, I. G. Petrovsky, N. S. Piskunov, Étude de l'équation de la diffusion avec croissance de la quantité de matière et son application à un probléme biologique, Bull. Univ. Moskov. Ser. Internat., 1 (1937), 1–25. |
[2] |
R. A. Fisher, The wave of advance of advantageous genes, Ann. Eugen., 7 (1937), 355–369. https://doi.org/10.1111/j.1469-1809.1937.tb02153.x doi: 10.1111/j.1469-1809.1937.tb02153.x
![]() |
[3] | M. Lewis, S. Petrovskii, J. Potts, The Mathematics Behind Biological Invasions, Springer, New York, (2016). https://doi.org/10.1007/978-3-319-32043-4 |
[4] |
K. Cuddington, A. Hastings, Invasive engineers, Ecol. Model., 178 (2004), 335–347. https://doi.org/10.1016/j.ecolmodel.2004.03.010 doi: 10.1016/j.ecolmodel.2004.03.010
![]() |
[5] |
C. G. Jones, J. H. Lawton, M. Shachak, Organisms as ecosystem engineers, Oikos, 69 (1994), 373–386. https://doi.org/10.1007/978-1-4612-4018-1_14 doi: 10.1007/978-1-4612-4018-1_14
![]() |
[6] |
C. G. Jones, J. H. Lawton, M. Shachak, Positive and negative effects of organisms as physical ecosystem engineers, Ecology, 78 (1997), 1946–1957. https://doi.org/10.1890/0012-9658(1997)078[1946:PANEOO]2.0.CO;2 doi: 10.1890/0012-9658(1997)078[1946:PANEOO]2.0.CO;2
![]() |
[7] |
A. Hasting, J. E. Byers, J. A. Crooks, K. Cuddington, C. G. Jones, J. G. Lambrinos, et al., Ecosystem engineering in space and time, Ecol. Lett., 10 (2007), 153–164. https://doi.org/10.1111/j.1461-0248.2006.00997.x doi: 10.1111/j.1461-0248.2006.00997.x
![]() |
[8] |
W. Ziebis, S. Forster, M. Huettel, B. Jørgensen, Complex burrows of the mud shrimp Callianassa truncata and their geochemical impact in the sea bed, Nature, 382 (1996), 619–622. https://doi.org/10.1038/382619a0 doi: 10.1038/382619a0
![]() |
[9] |
K. Cuddington, W. G. Wilson, A. Hastings, Ecosystem engineers: Feedback and population dynamics, Am. Nat., 173 (2009), 488–498. https://doi.org/10.1086/597216 doi: 10.1086/597216
![]() |
[10] |
F. Lutscher, J. Fink, Y. Zhu, Pushing the boundaries: Models for the spatial spread of ecosystem engineers, Bull. Math. Biol., 82 (2020), 138. https://doi.org/10.1007/s11538-020-00818-8 doi: 10.1007/s11538-020-00818-8
![]() |
[11] | F. Courchamp, L. Berec, J. Gascoigne, Allee effects in ecology and conservation, Oxford University Press, Oxford, UK (2008). https://doi.org/10.1093/acprof: oso/9780198570301.001.0001 |
[12] |
K. P. Hadeler, F. Rothe, Travelling fronts in nonlinear diffusion equations, J. Math. Biol., 2 (1975), 251–263. https://doi.org/10.1007/BF00277154 doi: 10.1007/BF00277154
![]() |
[13] |
Y. Du, Z. G. Lin, Spreading-vanishing dichotomy in the diffusive logistic model with a free boundary, SIAM J. Math. Anal., 42 (2010), 377–405. https://doi.org/10.1137/090771089 doi: 10.1137/090771089
![]() |
[14] | J. Stefan, Über die Theorie der Eisbildung insbesondere über die Eisbildung im Polarmeere, S. B. Wien Adak. Mat. Natur., 98 (1889), 965–983. |
[15] | L. Rubinstein, The Stefan Problem, American Mathematical Society, 1971. |
[16] |
G. Bunting, Y. Du, K. Krakowski, Spreading speed revisited: Analysis of a free boundary model, Netw. Heterog. Media, 7 (2012), 583–603. https://doi.org/10.3934/nhm.2012.7.583 doi: 10.3934/nhm.2012.7.583
![]() |
[17] |
Y. Du, Z. Guo, Spreading-vanishing dichotomy in a diffusive logistic model with a free boundary II, J. Diff. Equations, 250 (2011), 4336–4366. https://doi.org/10.1016/j.jde.2011.02.011 doi: 10.1016/j.jde.2011.02.011
![]() |
[18] |
Y. Du, Z. Guo, The Stefan problem for the Fisher-KPP equation, J. Diff. Equations, 253 (2012), 996–1035. http://dx.doi.org/10.1016/j.jde.2012.04.014 doi: 10.1016/j.jde.2012.04.014
![]() |
[19] |
Y. Du, A. G. Lin, The diffusive competition model with a free boundary: Invasion of a superior or inferior competitor, Discr. Cont. Dyn. Sys. B, 19 (2014), 3105–3132. https://doi.org/10.48550/arXiv.1303.0454 doi: 10.48550/arXiv.1303.0454
![]() |
[20] |
Y. Du, L. Wei, L. Zhou, Spreading in a shifting environment modeled by the diffusive logistic equation with a free boundary, J. Dyn. Diff. Equations, 30 (2018), 1389–1426. https://doi.org/10.1007/s10884-017-9614-2 doi: 10.1007/s10884-017-9614-2
![]() |
[21] |
Y. Du, J. Fang, N. Sun, A delay induced nonlocal free boundary problem, Math. Ann., 386 (2024), 2061–2106. https://doi.org/10.1007/s00208-022-02451-3 doi: 10.1007/s00208-022-02451-3
![]() |
[22] |
N. T. Fadai, Semi-infinite travelling waves arising in a general reaction-diffusion Stefan model, Nonlinearity, 34 (2021), 725–743. https://doi.org/10.1088/1361-6544/abd07b doi: 10.1088/1361-6544/abd07b
![]() |
[23] |
N. T. Fadai, Exact smooth and sharp-fronted traveling waves of reaction-diffusion equations with weak Allee effects, Appl. Math. Lett., 135 (2023), 108433. https://doi.org/10.1016/j.aml.2022.108433 doi: 10.1016/j.aml.2022.108433
![]() |
[24] |
C. Feng, M. A. Lewis, C. Wang, H. Wang, A Fisher-KPP model with a nonlocal weighted free boundary: Analysis of how habitat boundaries expand, balance or shrink, Bull. Math. Biol., 84 (2022), 34. https://doi.org/10.1007/s11538-022-00995-8 doi: 10.1007/s11538-022-00995-8
![]() |
[25] |
S. W. McCue, M. El-Hachem, M. J. Simpson, Traveling waves, blow-up, and extinction in the Fisher-Stefan model, Stud. Appl. Math., 148 (2022), 964–986. https://doi.org/10.1111/sapm.12465 doi: 10.1111/sapm.12465
![]() |
[26] | Y. Du, Propagation dynamics of the monostable reaction-diffusion equation with a new free boundary condition, Discr. Cont. Dyn. Sys. B, 44 (2024), 2524–2563. |
[27] |
G. Maciel, F. Lutscher, How individual movement response to habitat edges affects population persistence and spatial spread, Am. Nat., 182 (2013), 42–52. https://doi.org/10.1086/670661 doi: 10.1086/670661
![]() |
[28] |
J. S. MacDonald, F. Lutscher, Individual behavior at habitat edges may help populations persist in moving habitats, J. Math. Biol., 77 (2018), 2049–2077. https://doi.org/10.1007/s00285-018-1244-8 doi: 10.1007/s00285-018-1244-8
![]() |
[29] | M. Basiri, A Free Boundary Problem Modeling the Spread of Ecosystem Engineers, PhD thesis, University of Ottawa, 2023. http://dx.doi.org/10.20381/ruor-29166 |
[30] |
E. E. Crone, L. M. Brown, J. A. Hodgson, F. Lutscher, C. B. Schultz, Faster movement in habitat matrix promotes range shifts in heterogeneous landscapes, Ecology, 100 (2019), e02701. https://doi.org/10.1002/ecy.2701 doi: 10.1002/ecy.2701
![]() |
[31] |
M. Basiri, F. Lutscher, A. Moameni, Existence of solutions for a free boundary problem modeling the spread of ecosystem engineers, J. Nonlinear Sci., 31 (2021), 72. https://doi.org/10.1007/s00332-021-09725-1 doi: 10.1007/s00332-021-09725-1
![]() |
[32] |
H. Berestycki, L. Rossi, Reaction-diffusion equations for population dynamics with forced speed Ⅰ-The case of the whole space, Discr. Cont. Dyn. Sys. B, 21 (2008), 41–67. https://doi.org/10.3934/dcds.2008.21.41 doi: 10.3934/dcds.2008.21.41
![]() |
[33] |
J. S. MacDonald, Y. Bourgault, F. Lutscher, Moving habitat models: A numerical approach, Math. Biosci., 341 (2021), 108711. https://doi.org/10.1016/j.mbs.2021.108711 doi: 10.1016/j.mbs.2021.108711
![]() |
[34] |
B. Li, S. Bewick, J. Shang, W. F. Fagan, Persistence and spread of a species with a shifting habitat edge, SIAM J. Appl. Math., 74 (2014), 1397–1417. https://doi.org/10.1137/130938463 doi: 10.1137/130938463
![]() |
[35] |
J. B. Wang, W. T. Li, G. D. Dong, S. X. Qiao, Recent developments on spatial propagation for diffusion equations in shifting environments, Discr. Cont. Dyn. Sys. B, 27 (2022), 5101–5127. https://doi.org/10.3934/dcdsb.2021266 doi: 10.3934/dcdsb.2021266
![]() |
[36] | J. Bouhours, T. Giletti, Extinction and spreading of a species under the joint influence of climate change and a weak Allee effect: A two-patch model, preprint, arXiv: 1601.06589. |
[37] |
A. B. Potapov, M. A. Lewis, Climate and competition: The effect of moving range boundaries on habitat invasibility, Bull. Math. Biol., 66 (2004), 975–1008. https://doi.org/10.1016/j.bulm.2003.10.010 doi: 10.1016/j.bulm.2003.10.010
![]() |
[38] |
B. Li, G. Otto, Forced traveling waves in a reaction–diffusion equation with strong Allee effect and shifting habitat, Bull. Math. Biol., 85 (2023), 1–26. https://doi.org/10.1007/s11538-023-01221-9 doi: 10.1007/s11538-023-01221-9
![]() |
[39] | A. I. Volpert, V. A. Volpert, V. A. Volpert, Traveling wave solutions of parabolic systems, in Translations of Mathematical Monographs, American Mathematical Society, (1994), 448. |
[40] |
A. Hastings, K. Cuddington, K. F. Davies, C. J. Dugaw, S. Elmendorf, A. Freestone, et al., The spatial spread of invasions: new developments in theory and evidence, Ecol. Lett., 8 (2005), 91–101. https://doi.org/10.1111/j.1461-0248.2004.00687.x doi: 10.1111/j.1461-0248.2004.00687.x
![]() |